[1
]Department of Ecology and Biogeography Nicolaus Copernicus University Toruń Poland
[2
]Faculty of Science University of the Ryukyus Nishihara Japan
[3
]University MuseumUniversity of the Ryukyus Nishihara Japan
[4
]Royal Botanic Gardens Kew, Richmond United Kingdom
[5
]Marine and Terrestrial Field Ecology Tropical Biosphere Research Center University
of the Ryukyus Nishihara Japan